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This Project Gutenberg was produced by Marilynda Fraser-Cunliffe, LN Yaddanapudi and the Online Distributed Proofreading Team at http://www.pgdp.net

[i]

CONTRIBUTIONS TO
THE THEORY OF
NATURAL SELECTION.
A Series of Essays.
BY
ALFRED RUSSEL WALLACE,
AUTHOR OF
“THE MALAY ARCHIPELAGO,” ETC., ETC.
SECOND EDITION, WITH CORRECTIONS AND ADDITIONS.

New York:
MACMILLAN AND CO.
1871.

[ii]

  1. On the Law which has regulated the introduction of New Species.
    • Geographical distribution dependent on Geologic Changes
    • A Law deduced from well-known Geographical and Geological facts
    • The form of a true system of Classification determined by this Law
    • Geographical Distribution of Organisms
    • Geological Distribution of the forms of Life
    • High Organization of very ancient Animals consistent with this Law
    • Objections to Forbes’ Theory of Polarity
    • Rudimentary Organs
    • Conclusion
  2. On the Tendency of Varieties to depart indefinitely from the Original Type.
    • Instability of Varieties supposed to prove the permanent distinctness of Species
    • The Struggle for Existence
    • The Law of Population of Species
    • The Abundance or Rarity of a Species dependent upon its more or less perfect Adaptation to the Conditions of Existence
    • Useful Variations will tend to Increase, useless or hurtful Variations to Diminish
    • Superior Varieties will ultimately extirpate the Original Species
    • The Partial Reversion of Domesticated Varieties explained
    • Lamarck’s Hypothesis very different from that now advanced
    • Conclusion
  3. Mimicry, and other Protective Resemblances among Animals.
    • Test of true and false Theories
    • Importance of the Principle of Utility
    • Popular Theories of Colour in Animals
    • Importance of Concealment as influencing Colour
    • Special modifications of Colour
    • Theory of Protective Colouring
    • Objection that Colour as being dangerous should not exist in Nature
    • Mimicry
    • Mimicry among Lepidoptera
    • Lepidoptera mimicking other Insects
    • Mimicry among Beetles
    • Beetles mimicking other Insects
    • Insects mimicking Species of other Orders
    • Cases of Mimicry among the Vertebrata
    • Mimicry among Snakes
    • Mimicry among Birds
    • Mimicry among Mammals[xiv]
    • Objections to Mr. Bates’ Theory of Mimicry
    • Mimicry by Female Insects only
    • Cause of the dull Colours of Female Birds
    • Use of the gaudy Colours of many Caterpillars
    • Summary
    • General deductions as to Colour in Nature
    • Conclusion
  4. The Malayan Papilionidæ, or Swallow-tailed Butterflies, as illustrative of the Theory of Natural Selection.
    • Special value of the Diurnal Lepidoptera for inquiries of this Nature
    • Question of the rank of the Papilionidæ
    • Distribution of the Papilionidæ
    • Definition of the word Species
    • Laws and Modes of Variation
      • Simple Variability
      • Polymorphism or Dimorphism
      • Local form or variety
      • Co-existing Variety
      • Race or Subspecies
      • Species
    • Variation as specially influenced by Locality
      • Local Variation of Size
      • Local Variation of Form
      • Local Variations of Colour
    • Remarks on the facts of Local Variation
    • Mimicry
    • Concluding Remarks on Variation in Lepidoptera
    • Arrangement and Geographical Distribution of the Malayan Papilionidæ
      • Arrangement
      • Geographical Distribution
    • Range of the Groups of Malayan Papilionidæ
    • Remarkable peculiarities of the island of Celebes
    • Concluding Remarks
  5. On Instinct in Man and Animals.
    • How Instinct may be best Studied
    • Definition of Instinct
    • Does Man possess Instincts?
    • How Indians travel through unknown and trackless Forests
  6. The Philosophy of Birds’ Nests.
    • Instinct or Reason in the Construction of Birds’ Nests
    • Do Men build by Reason or by Imitation?
    • Why does each Bird build a peculiar kind of Nest?
    • How do young Birds learn to build their first Nest?
    • Do Birds sing by Instinct or by Imitation?
    • How young Birds may learn to build Nests.
    • Man’s Works mainly Imitative
    • Birds do Alter and Improve their Nests when altered conditions require it
    • Conclusion
  7. A Theory of Birds’ Nests; showing the relation of certain differences of colour in female birds to their mode of nidification.
    • Changed Conditions and persistent Habits as influencing Nidification
    • Classification of Nests
    • Sexual differences of Colour in Birds
    • The Law which connects the Colours of Female Birds with the mode of Nidification
    • What the Facts Teach us
    • Colour more variable than Structure or Habits, and therefore the Character which has generally been modified
    • Exceptional cases confirmatory of the above Explanation
    • Real or apparent exceptions to the Law stated at p. 240
    • Various modes of Protection of Animals
    • Females of some groups require and obtain more Protection than the Males
    • Conclusion
  8. Creation by Law.
    • Laws from which the Origin of Species may be deduced
    • Mr. Darwin’s Metaphors liable to Misconception
    • A case of Orchis-structure explained by Natural Selection
    • Adaptation brought about by General Laws
    • Beauty in Nature
    • How new Forms are produced by Variation and Selection
    • The Objection that there are Limits to Variation
    • Objection to the argument from Classification
    • The Times on Natural Selection
    • Intermediate or generalized forms of Extinct Animals an indication of Transmutation or Development
    • Conclusion
    • A Demonstration of the Origin of Species
  9. The Development of Human Races under the Law of Natural Selection.
    • Wide difference of Opinion as to Man’s Origin
    • Outline of the Theory of Natural Selection
    • Different effects of Natural Selection on Animals and on Man
    • Influence of External Nature in the development of the Human Mind
    • Extinction of Lower Races
    • The Origin of the Races of Man
    • The Bearing of these views on the Antiquity of Man
    • Their Bearing on the Dignity and Supremacy of Man
    • Their Bearing on the future Development of Man
    • Summary
    • Conclusion
  10. The Limits of Natural Selection as applied to Man.
    • What Natural Selection can Not do
    • The Brain of the Savage shown to be Larger than he Needs it to be
      • Size of Brain an important Element of Mental Power
      • Comparison of the Brains of Man and of Anthropoid Apes
      • Range of intellectual power in Man
      • Intellect of Savages and of Animals compared
    • The use of the Hairy Covering of Mammalia
    • The Constant absence of Hair from certain parts of Man’s body a remarkable Phenomenon
    • Savage Man feels the want of this Hairy Covering
    • Man’s Naked Skin could not have been produced by Natural Selection
    • Feet and Hands of Man considered as Difficulties on the Theory of Natural Selection
    • The Origin of Some of Man’s Mental Faculties, by the preservation of Useful Variations, not possible
    • Difficulty as to the Origin of the Moral Sense
    • Summary of the Argument as to the Insufficiency of Natural Selection to account for the Development of Man
    • The Origin of Consciousness
    • The Nature of Matter
      • Matter is Force
      • All Force is probably Will-force
    • Conclusion


[iii]

PREFACE.

The present volume consists of essays which I have contributed to various periodicals, or read before scientific societies during the last fifteen years, with others now printed for the first time. The two first of the series are printed without alteration, because, having gained me the reputation of being an independent originator of the theory of “natural selection,” they may be considered to have some historical value. I have added to them one or two very short explanatory notes, and have given headings to subjects, to make them uniform with the rest of the book. The other essays have been carefully corrected, often considerably enlarged, and in some cases almost rewritten, so as to express more fully and more clearly the views which I hold at the present time; and as most of them originally appeared in publications which have a very limited circulation, I believe that the larger portion of this volume will be new to many of my friends and to most of my readers.

I now wish to say a few words on the reasons which have led me to publish this work. The second essay, especially when taken in connection with the first, contains an outline sketch of the theory of the origin of species (by means of what was afterwards termed by Mr. Darwin—“natural selection,”) as conceived [iv] by me before I had the least notion of the scope and nature of Mr. Darwin’s labours. They were published in a way not likely to attract the attention of any but working naturalists, and I feel sure that many who have heard of them, have never had the opportunity of ascertaining how much or how little they really contain. It therefore happens, that, while some writers give me more credit than I deserve, others may very naturally class me with Dr. Wells and Mr. Patrick Matthew, who, as Mr. Darwin has shown in the historical sketch given in the 4th and 5th Editions of the “Origin of Species,” certainly propounded the fundamental principle of “natural selection” before himself, but who made no further use of that principle, and failed to see its wide and immensely important applications.

The present work will, I venture to think, prove, that I both saw at the time the value and scope of the law which I had discovered, and have since been able to apply it to some purpose in a few original lines of investigation. But here my claims cease. I have felt all my life, and I still feel, the most sincere satisfaction that Mr. Darwin had been at work long before me, and that it was not left for me to attempt to write “The Origin of Species.” I have long since measured my own strength, and know well that it would be quite unequal to that task. Far abler men than myself may confess, that they have not that untiring patience in accumulating, and that wonderful skill in using, large masses of facts of the [v] most varied kind,—that wide and accurate physiological knowledge,—that acuteness in devising and skill in carrying out experiments,—and that admirable style of composition, at once clear, persuasive and judicial,—qualities, which in their harmonious combination mark out Mr. Darwin as the man, perhaps of all men now living, best fitted for the great work he has undertaken and accomplished.

My own more limited powers have, it is true, enabled me now and then to seize on some conspicuous group of unappropriated facts, and to search out some generalization which might bring them under the reign of known law; but they are not suited to that more scientific and more laborious process of elaborate induction, which in Mr. Darwin’s hands has led to such brilliant results.

Another reason which has led me to publish this volume at the present time is, that there are some important points on which I differ from Mr. Darwin, and I wish to put my opinions on record in an easily accessible form, before the publication of his new work, (already announced,) in which I believe most of these disputed questions will be fully discussed.

I will now give the date and mode of publication of each of the essays in this volume, as well as the amount of alteration they have undergone.

I.—On the Law which has Regulated the Introduction of New Species.

First published in the “Annals and Magazine of [vi] Natural History,” September, 1855. Reprinted without alteration of the text.

II.—On the Tendency of Varieties to Depart indefinitely from the Original Type.

First published in the “Journal of the Proceedings of the Linnæan Society,” August, 1858. Reprinted without alteration of the text, except one or two grammatical emendations.

III.—Mimicry and other Protective Resemblances among Animals.

First published in the “Westminster Review,” July, 1867. Reprinted with a few corrections and some important additions, among which I may especially mention Mr. Jenner Weir’s observations and experiments on the colours of the caterpillars eaten or rejected by birds.

IV.—The Malayan Papilionidæ, Or Swallow-Tailed Butterflies, as Illustrative of the Theory of Natural Selection.

First published in the “Transactions of the Linnæan Society,” Vol. XXV. (read March, 1864), under the title, “On the Phenomena of Variation and Geographical Distribution, as illustrated by the Papilionidæ of the Malayan Region.”

The introductory part of this essay is now reprinted, omitting tables, references to plates, &c., with some additions, and several corrections. Owing to the publication [vii] of Dr. Felder’s “Voyage of the Novara” (Lepidoptera) in the interval between the reading of my paper and its publication, several of my new species must have their names changed for those given to them by Dr. Felder, and this will explain the want of agreement in some cases between the names used in this volume and those of the original paper.

V.—On Instinct in Man and Animals.

Not previously published.

VI.—The Philosophy of Birds’ Nests.

First published in the “Intellectual Observer,” July, 1867. Reprinted with considerable emendations and additions.

VII.—A Theory of Birds’ Nests; Showing the relation of certain differences Of Colour in Birds To their mode of Nidification.

First published in the “Journal of Travel and Natural History” (No. 2), 1868. Now reprinted with considerable emendations and additions, by which I have endeavoured more clearly to express, and more fully to illustrate, my meaning in those parts which have been misunderstood by my critics.

VIII.—Creation by Law.

First published in the “Quarterly Journal of Science,” October, 1867. Now reprinted with a few alterations and additions.

[viii]

IX.—The Development of Human Races under the Law of Natural Selection.

First published in the “Anthropological Review,” May, 1864. Now reprinted with a few important alterations and additions. I had intended to have considerably extended this essay, but on attempting it I found that I should probably weaken the effect without adding much to the argument. I have therefore preferred to leave it as it was first written, with the exception of a few ill-considered passages which never fully expressed my meaning. As it now stands, I believe it contains the enunciation of an important truth.

X.—The Limits of Natural Selection as applied to Man.

This is the further development of a few sentences at the end of an article on “Geological Time and the Origin of Species,” which appeared in the “Quarterly Review,” for April, 1869. I have here ventured to touch on a class of problems which are usually considered to be beyond the boundaries of science, but which, I believe, will one day be brought within her domain.


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